Page - 66 - in The Origin of Species

66 ORIGIN OF SPECIES Many of the cases of strongly-marked varieties or doubtful species well deserve consideration ; for several interesting lines of argument, from geographical distribution, analogical variation, hybridism, &c., have been brought to bear in the attempt to determine their rank ; but space does not here per- mit me to discuss them. Close investigation, in many cases, v^^ill no doubt bring naturalists to agree how to rank doubt- ful forms. Yet it must be confessed that it is in the best known countries that we find the greatest number of them. I have been struck with the fact, that if any animal or plant in a state of nature be highly useful to man, or from any cause closely attracts his attention, varieties of it will almost universally be found recorded. These varieties, moreover, will often be ranked by some authors as species. Look at the common oak, how closely it has been studied; yet a German author makes more than a dozen species out of forms, which are almost universally considered by other botanists to be varieties ; and in this country the highest botanical authori- ties and practical men can be quoted to show that the sessile and pedunculated oaks are either good and distinct species or mere varieties. I may here allude to a remarkable memoir lately published by A. de CandoUe, on the oaks of the whole world. No one ever had more ample materials for the discrimination of the species, or could have worked on them with more zeal and sagacity. He first gives in detail all the many points of struc- ture which vary in the several species, and estimates numeri- cally the relative frequency of the variations. He specifies above a dozen characters which may be found varying even on the same branch, sometimes according to age or develop- ment, sometimes without any assignable reason. Such char- acters are not of course of specific value, but they are, as Asa Gray has remarked in commenting on this memoir, such as generally enter into specific definitions. De Candolle then goes on to say that he gives the rank of species to the forms that differ by characters never varying on the same tree, and never found connected by intermediate states. After this discussion, the result of so much labour, he emphatically re- marks: "They are mistaken, who repeat that the greater part of our species are clearly limited, and that the doubtful