Page - 460 - in The Origin of Species

460 ORIGIN OF SPECIES In confirmation of this view, let us glance at the classifi- cation of varieties, which are known or believed to be descended from a single species. These are grouped under the species, with the sub-varieties under the varieties; and in some cases, as with the domestic pigeon, with several other grades of difference. Nearly the same rules are fol- lowed as in classifying species. Authors have insisted on the necessity of arranging varieties on a natural instead of an artificial system; we are cautioned, for instance, not to class two varieties of the pine-apple together, merely because their fruit, though the most important part, happens to be nearly identical ; no one puts the Swedish and common turnip together, though the esculent and thickened stems are so similar. Whatever part is found to be most constant, is used in classing varieties; thus the great agriculturist Marshall says the horns are very useful for this purpose with cattle, because they are less variable than the shape or colour of the body, &c. ; whereas with sheep the horns are much less serv- iceable, because less constant. In classing varieties, I apprehend that if we had a real pedigree, a genealogical classification would be universally preferred; and it has been attempted in some cases. For we might feel sure, whether there had been more or less modification, that the principle of inheritance would keep the forms together which were allied in the greatest number of points. In tumbler pigeons, though some of the sub-varieties differ in the important character of the length of the beak, yet all are kept together from having the common habit of tumbling; but the short- faced breed has nearly or quite lost this habit; nevertheless, without any thought on the subject, these tumblers are kept in the same group, because allied in blood and alike in some other respects. With species in a state of nature, every naturalist has in fact brought descent into his classification; for he includes in his lowest grade, that of species, the two sexes ; and how enormously these sometimes differ in the most important characters, is known to every naturalist : scarcely a single fact can be predicated in common of the adult males and hermaphrodites of certain cirripedes, and yet no one dreams of separating them. As soon as the three Orchidean forms,